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Here’s a list of  my published research. Depending on your time and interest, please click on the links beside each to view in your preferred format.

2021 & In press

Rafiq, K., Appleby, R.G., Edgar, J.P., Radford, C., Smith, B.P., Jordan, N.R., Dexter, C.E., Jones, D.N., Blacker, A.R.F. and Cochrane, M. (2021). WildWID: an open‐source active RFID system for wildlife research. Methods in Ecology and Evolution. https://doi.org/10.1111/2041-210X.13651

Rafiq, K., Pitcher, B.J., Cornelsen, K., Hansen, K.W., King, A.J., Appleby, R.G., Abrahms, B. & Jordan, N.R. (2021). Animal-borne technologies in wildlife research and conservation. Conservation Technology, p.105. (View)

Alting, B. F., Bennitt, E., Golabek, K. A., Pitcher, B. J., McNutt, J. W., Wilson, A. M., Bates, H. & Jordan, N. R. (2021). The characteristics and consequences of African wild dog (Lycaon pictus) den site selection. Behavioral Ecology and Sociobiology, 75(7), 1-14. https://rdcu.be/cpdz3

Downey, H., Amano, T., Cadotte, M., Cook, C. N., Cooke, S. J., Haddaway, N. R., … Jordan, N. R., … & Sutherland, W. J. (2021). Training future generations to deliver evidence‐based conservation and ecosystem management. Ecological Solutions and Evidence, 2(1), e12032. https://doi.org/10.1002/2688-8319.12032

Buesching, C.D. & Jordan, N.R. (In press) Small carnivore latrines in space and time (with special emphasis on badgers and mongooses). In: Small Carnivores in Space and Time. Ed. E Do Linh San. Cambridge University Press, Cambridge UK. Part II, Chapter 11.

Smith BP, Jaques NB, Appleby RG, Morris S & Jordan NR. (2021). Automated shepherds: responses of captive dingoes to sound and an inflatable, moving effigy. Pacific Conservation Biology https://doi.org/10.1071/PC20022


Witt RR, Beranek CT, Howell LG, Ryan SA, Clulow J, Jordan NR, et al. (2020) Real-time drone derived thermal imagery outperforms traditional survey methods for an arboreal forest mammal. PLoS ONE 15(11): e0242204. https://doi.org/10.1371/journal.pone.0242204

Vitale JD, Jordan NR, Gilfillan GD, McNutt JW, Reader T. (2020). Spatial and seasonal patterns of communal latrine use by spotted hyenas (Crocuta crocuta) reflect a seasonal resource defense strategy. Behavioral Ecology and Sociobiology 74 (10), 1-14

Smith BP, Appleby RG & Jordan NR. (2020). Co-existing with dingoes: Challenges and solutions to implementing non-lethal management. Australian Zoologist https://doi.org/10.7882/AZ.2020.024

Radford, C., McNutt, J.W., Rogers, T., Maslen, B. & Jordan, N.R. (2020). Artificial eyespots on cattle reduce predation by large carnivores. Communications Biology 3, 430. https://doi.org/10.1038/s42003-020-01156-0

Jordan, N.R., Smith, B.P., Appleby, R.G., van Eeden, L.M. & Webster, H.S. (2020). Addressing inequality and intolerance in human-wildlife coexistence. Conservation Biology. https://doi.org/10.1111/cobi.13471

Adams, T.S. Mwezi, I. & Jordan, N.R. (2020). Panic at the disco: Solar-powered strobe lights reduce field incursions by African elephants, in the Chobe District, Botswana. Oryx. https://doi.org/10.1017/S0030605319001182

Rafiq, K., Hayward, M.W., Wilson, A.M., Meloro, C., Jordan, N.R., Wich, S.A., McNutt, J.W. and Golabek, K.A. (2020). Spatial and temporal overlaps between leopards (Panthera pardus) and their competitors in the African large predator guild. Journal of Zoology. Full Article; Short & Tweet.

Jordan, N.R. (2020).  Making Sense of Scents: Olfactory Perception in Animals. Box A3, In Zoo Animal Learning and Training, First Edition. Edited by Melfi, V.A., Dorey, N.R. & Ward, S.J. Wiley & Sons Ltd. Buy the Book.

Rafiq, K., Jordan, N.R., Meloro, C., Wilson, A., Hayward, M & McNutt, J.W. (2020) Optimal scent marking strategies of a solitary carnivore: boundary and road scent marking in the leopard. Animal Behaviour 161, 115-126. Full Article; Short & Tweet.


Buesching, C. D., & Jordan, N. (2019). The social function of latrines: A hypothesis-driven research approach. In Chemical Signals in Vertebrates 14 (pp. 94-103). Springer, Cham. Full Article; Short & Tweet.

Rafiq, K., Jordan, N.R., Wilson, A.M., McNutt, J.W., Hayward, M.W., Meloro, C., Golabek, K. (2019) Spatio-temporal factors impacting encounter occurrences between leopards and other large African predators. Journal of Zoology 310, 191-200. Full Article; Short & Tweet.

Brink, H., Purcell, B.V., Letnic, M., Webster, H.S., Appleby, R.G. & Jordan, N.R. (2019). The contrasting economics of canid pets and pests in Australia: harnessing humanity’s affiliation for domestic dogs to conserve their wild counterparts. Wildlife Research 46, 365-377. Full Article; Article in The Conversation.

Rafiq, K., Appleby, R.G., Edgar, J.P., Jordan, N.R., Blacker, A.R., Dexter, C.E., Jones, D.N. and Cochrane, M. (2019). OpenDropOff: an open-source, low-cost drop-off unit for animal borne devices. Methods in Ecology and Evolution 10, 1517-1522. Full Article; Short & Tweet.

Chisholm, S., Stein, A.B., Jordan, N.R., Hubel, T.M., Shawe-Taylor, J., Fearn, T., McNutt, J.W., Wilson, A.M. & Hailes, S. (2019) Parsimonious Test of Dynamic Interaction. Ecology and Evolution 9, 1654-1664. Full Article.

Kusler, A., Jordan, N.R., McNutt, J. & Broekhuis, F. (2019). Cheetah marking trees: distribution, visitation, and behavior. African Journal of Ecology 57, 419-422. Full Article; Short & Tweet.

Bryant, B. Pittard, S., Jordan, N.R. & McMahon, C.R. (2019). The chemical capture of wild swamp buffalo (Bubalus bubalis) in tropical Australia using thiafentanil, etorphine and azaperone combinations. Australian Veterinary Journal 97, 33-38. Full Article.


McNutt, J.W., Stein, A.B., Boggs-McNutt, L. & Jordan, N.R. (2018). Living on the edge: Characteristics of human-wildlife conflict in a traditional livestock community in Botswana. Wildlife Research 44(7) 546-557. Full Article


Torrents Ticó, M., Rich, L., McNutt, J.W., Nnthomiwa, M., Mothala, M., Motsamai, G. & Jordan, N.R. (2017). On the right track? Comparing concurrent spoor and camera-trap surveys in Botswana. African Journal of Wildlife Research 47(2):128-137; DOI: https://doi.org/10.3957/056.047.0128. Full article; Short & tweet.

Walker, R.H., King, A.J., McNutt, J.W. & Jordan, N.R. (2017). Sneeze to leave: African wild dogs (Lycaon pictus) use variable quorum thresholds mediated by sneezes in collective decisions. Proceedings of the Royal Society, B. 284, 20170347; DOI: 10.1098/rspb.2017.0347. Full article;  Blog;  Press; Short & tweet.

Jordan, N.R., O’Riain, J., Balmforth, Z., Martins, V., Page-Nicholson, S., Do Linh San, E. (2017). A conservation assessment of Suricata suricatta. In Child MF, Roxburgh L, Do Linh San E, Raimondo D, Davies-Mostert HT, editors. The Red List of Mammals of South Africa, Swaziland and Lesotho. South African National Biodiversity Institute and Endangered Wildlife Trust, South Africa.

Jordan, N.R., Buse, C., Wilson, A.M., Golabek, K.A., Apps, P.J., Lowe, J.C., Van der Weyde, L.K. & McNutt, J.W. (2017). Dynamics of direct inter-pack encounters in endangered African wild dogs. Behavioral Ecology and Sociobiology, 71, 115. Full article;  Short & tweet.

Jackson, C.R., Groom, R., Jordan, N.R. & McNutt, J.W. (2017). The effect of relatedness and pack size on territory overlap in African wild dogs. Movement Ecology 5.1 (2017): 10. Full article; Short & tweet.

Dewhirst, O.P., Roskilly, K., Hubel, T.Y., Jordan, N.R., Golabek, K.A., McNutt, J.W. & Wilson, A.M. (2017). An exploratory clustering approach for extracting stride parameters from tracking collars on free ranging wild animals. Journal of Experimental Biology. doi: 10.1242/jeb.146035. Full article.


Abrahms, B., Sawyer, S. C., Jordan, N. R., McNutt, J. W., Wilson, A. M., Brashares, J. S. (2016), Does wildlife resource selection accurately inform corridor conservation? Journal of Applied Ecology. doi: 10.1111/1365-2664.12714. Full article; short & tweet.

Hubel, T.Y., Myatt, J.P., Jordan, N.R., Dewhirst, O.P., McNutt, J.W. & Wilson A.M. (2016) Additive opportunistic capture explains group hunting benefits in African wild dogs. Nature Communications. DOI: 10.1038/ncomms11033. Full article; blog & press; short & tweet.

Hubel, T.Y., Myatt, J.P., Jordan, N.R., Dewhirst, O.P., McNutt, J.W. & Wilson A.M. (2016) Energy cost and return for hunting in African wild dogs and cheetahs. Nature Communications. DOI: 10.1038/ncomms11034. Full article; blog & press; short & tweet.

Jordan, N.R., Apps, P.J., Golabek, K.A. & McNutt, J.W. (2016) Pair-specific scents in African wild dogs, Lycaon pictus, and an example of a potential method to identify signals within complex mixtures. Chapter 30, In: Chemical Signals in Vertebrates 13. Ed. BA Schulte, T Goodwin & M Ferkin. New York: D. Kluwer Academic/Plenum Publishers. Full article; short & tweet.

Abrahms, B., Jordan, N.R., Golabek, K.A., McNutt, J.W., Wilson, A.M. & Brashares, J.S. (2016) Lessons from integrating behaviour and resource selection: activity-specific responses of African wild dogs to roads. Animal Conservation. DOI: 10.1111/acv.12235. Full article; short & tweet.


Pomilia, M.A., McNutt, J.W. & Jordan, N.R. (2015) Ecological predictors of African wild dog ranging patterns in northern Botswana. Journal of Mammalogy. DOI: http://dx.doi.org/10.1093/jmammal/gyv130. Full article; short & tweet.

Abrahms, B., Sawyer, S.C, Jordan, N.R., McNutt, J.W., Wilson, A.M. & Brashares, J. “Do estimates of landscape resistance reflect movement behavior?”  [abstract]. In: Proceedings of the 27th International Congress for Conservation Biology and 4th European Congress for Conservation Biology. Visconti P., Game E., Mathevet R., Wilkerson M. editors. Montpellier 2-6 August 2015. p. 5. SCB; 2015. doi: 10.13140/RG.2.1.4233.4489/1. Abstract.

Jordan, N.R. & Do Linh San, E. 2015. Suricata suricatta. The IUCN Red List of Threatened Species 2015: e.T41624A45209377. http://dx.doi.org/10.2305/IUCN.UK.2015-4.RLTS.T41624A45209377.en. Full article.


Jordan, N.R., Apps, P.J., Golabek, K.A. & McNutt, J.W. 2014. Top marks for top dogs: Tandem marking and pair-bond advertisement in African wild dogs (Lycaon pictus). Animal Behaviour 88, 211-217. Full article.

 Apps, P., Mmualefe, L., Jordan, N.R., Golabek, K.A. & McNutt, J.W. 2014. The “tomcat compound” 3-mercapto-3-methylbutanol occurs in the urine of free-ranging leopards but not in African lions or cheetahs. Biochemical Systematics and Ecology 53, 17-19.Full article.


 Jordan, N.R., Golabek, K.A., Apps, P.J., Gilfillan, G.D. & McNutt, J.W. 2013. Scent-mark identification and scent-marking behaviour in African wild dogs (Lycaon pictus). Ethology 119, 1-9. Full article.


Nichols, H., Jordan, N.R., Jamie, G., Cant, M. & Hoffman, J. 2012. Fine-scale spatiotemporal patterns of genetic variation reflect budding dispersal coupled with strong natal philopatry in a cooperatively breeding mammal Molecular Ecology 21, 5348-5362.Full article.

 Jordan, N.R., Messenger, J.E., Turner, P., Birks, J.D.S., Croose, E. & O’Reilly, C. 2012. Molecular comparison of historical and contemporary pine marten (Martes martes) populations in the British Isles: evidence of differing origins and fates. Conservation Genetics 13, 1195- 1212. Full article.

Powell, R.A., Lewis, J C., Slough, B. G., Brainerd, S.M., Jordan, N.R., Abramov, A.V., Monakhov, V.,  Zollner, P. &  Murakami, T. 2012. Evaluating translocations of martens, sables and fishers: Testing model predictions with field data. In: Biology and conservation of martens, sables and fishers: a new synthesis. Ed. KB Aubry, WJ Zielinski, MG Raphael, G Proulx, SW Buskirk. Cornell University Press. Chapter 6, pp. 93-137.


Jordan, N.R. 2011. Strategy for restoring the pine marten to England and Wales. Report published by The Vincent Wildlife Trust.

 Jordan, N.R., Manser, M.B., Mwanguhya, F., Kyabulima, S., Rüedi, P. & Cant, M.A. 2011. Scent marking in wild banded mongooses: 1. Sex specific scents and over-marking. Animal Behaviour 81, 31-42. Full article.

Jordan, N.R., Mwanguhya, F., Furrer, R.D., Kyabulima, S., Rüedi, P. & Cant, M.A. 2011. Scent marking in wild banded mongooses: 2. Intrasexual over-marking and competition between males. Animal Behaviour 81, 43-50. Full article.

Jordan, N.R., Mwanguhya, F., Kyabulima, S., Rüedi, P. Hodge, S.J. & Cant, M.A. 2011. Scent marking in wild banded mongooses: 3. Intrasexual over-marking in females. Animal Behaviour 81, 51-60. Full article.


Jordan, N.R., Mwanguhya, F., Kyabulima, S., Rüedi, P. & Cant, M.A. 2010. Scent marking within and between groups of wild banded mongooses. Journal of Zoology 280, 72-83. Full article.


Jordan, N.R. 2009. Scent communication in wild banded mongooses. PhD Thesis. University of Cambridge.

Golabek, K.A., Jordan, N.R. & Clutton-Brock, T.H. 2008. Radiocollars do not affect the survival or foraging behaviour of wild meerkats. Journal of Zoology 274, 248-253. Full article.

 Jordan, N.R. 2007. Scent marking investment is determined by sex and breeding status in meerkats. Animal Behaviour 74, 531-540. Full article.

 Jordan, N.R., Cherry, M.I. & Manser, M.B. 2007. Latrine distribution and patterns of use by wild meerkats: implications for territory and mate defence. Animal Behaviour 73, 613-622. Full article.

Clutton-Brock, T.H., Hodge, S.J., Spong, G., Russell, A.F., Jordan, N.R., Bennett, N.C., Sharpe, L.L. & Manser, M.B. 2006. Intrasexual competition and sexual selection in cooperative mammals. Nature 444, 1065-1068. Full article.

Russell, A.F., Young, A.J., Spong, G., Jordan, N.R. & Clutton-Brock, T.H. 2006. Helpers increase the reproductive potential of offspring in cooperative meerkats. Proceedings of the Royal Society B. 274, 513-520. Full article.

Carlson, A.A., Russell, A.F., Young, A.J., Jordan, N.R, McNeilly, A.S., Parlow, A.F. & Clutton-Brock, T.H. 2006. Elevated prolactin levels immediately precede decisions to babysit by male meerkat helpers. Hormones and Behaviour 50, 94-100. Full article.

Carlson, A.A., Manser, M.B., Young, A.J., Russell, A.F., Jordan, N.R, McNeilly, A.S. & Clutton-Brock, T H. 2006. Cortisol levels are positively associated with pup-feeding rates in male meerkats. Proceedings of the Royal Society B. 273, 571-577. Full article.

Jordan, N.R. 2005. Meerkat latrines: cooperation, competition and discrimination. MSc Thesis. University of Stellenbosch.

Clutton-Brock, T.H., Russell, A.F., Sharpe, L.L. & Jordan, N.R. 2005. ‘False-feeding’ and aggression in meerkat societies. Animal Behaviour 69, 1273-1284. Full article.

Russell, A.F., Carlson, A.A., McIlrath, G.M., Jordan, N.R & Clutton-Brock, T.H. 2004. Adaptive size modification by dominant female meerkats. Evolution 58, 1600-1607. Full article.

Plowman, A.B., Jordan, N.R, Anderson, N., Condon, E. & Fraser, O. 2004. Welfare implications of captive primate population management: behavioural and psycho-social effects of female-based contraception, oestrous and male removal in hamadryas baboons (Papio hamadryas). Applied Animal Behaviour Science 90, 155-165. Full article.